Bonobos are classified as Pan paniscus. They are the closest living relatives of chimpanzees. These two species diverged from common ancestors around 1.8 million years ago. Today bonobos live within a geographic range south of the Congo River.
The footprints of extinct lineages are the closest we have to a fossil record of the African apes.
A 33,000-year-old case of an amputated leg prompts comparisons to earlier Neandertal instances of amputation.